These are some representative publications that give an overview of the kinds of research we’re pursuing in the lab. A complete list of Elsa Cleland’s publications can be found here.

*UCSD graduate student, ‡ UCSD Postdoctoral Scholar


*Waterton, J. Mazer, S. J. and E. E. Cleland (2023) When the neighborhood matters: contextual selection on emergence time in native and exotic California grasses Evolution,

*Ryan, E.M. and E. E. Cleland. (2021) Clinal variation in phenological traits associated with drought response of California poppy across its native range. Climate Change Ecology 2: 100021

*Waterton, J. & E.E. Cleland. (2021) Mammalian herbivores weaken selection for early emergence in competition. Evolution Letters, doi:10.1002/evl3.222

*Esch, EH, D. Lipson & E.E. Cleland (2019) Invasion alters ecosystem response to drought via increased phenological sensitivity. Ecology 100: e02802

*Esch E.H., *A.C. Ashbacher, *C.W. Kopp, & E.E. Cleland (2018) Competition reverses the response of shrub seedling mortality and growth along a soil moisture gradient. Journal of Ecology 106: 2096-2108.

Cleland, E.E., *E.E. Esch and *J.G. McKinney (2015) Priority effects vary with species identity and origin in an experiment varying the timing of seed arrival. Oikos 124: 33-40.

*Kopp, C. & E.E. Cleland. (2014) Shifts in elevational range limits and alpine plant species abundances observed over nearly five decades in a western North America mountain range. Journal of Vegetation Science 25: 135-146.

Cleland, E.E., J.M. Allen, T.M. Crimmins, J.A. Dunne, S. Pau, S. Travers, E.S. Zavaleta, and ‡ E.M. Wolkovich (2012) Phenological tracking enables positive species responses to climate change. Ecology 93:1765–1771.

‡ Wolkovich, E.M., B.I. Cook, J.M. Allen,T.M. Crimmins, J.L. Betancourt, S. Travers, S. Pau, J. Regetz, T.J. Davies, N.J.B. Kraft, T.R. Ault, K. Bolmgren, S.J. Mazer, G.J. McCabe, B.J. McGill, C. Parmesan, N. Salamin, M.D. Schwartz & E.E. Cleland (2012) Warming experiments underpredict plant phenological responses to climate change. Nature 485:494-7.